:root {
  --bg: #f6f4ee;
  --panel: #fffdf8;
  --panel-strong: #f1ede2;
  --border: #ddd4c5;
  --text: #1f1d17;
  --muted: #666052;
  --accent: #5c4b1f;
  --accent-soft: #d9caa5;
  --shadow: 0 10px 30px rgba(55, 44, 25, 0.08);
  --radius: 18px;
  --radius-small: 12px;
  --max: 1400px;
  --font: Inter, ui-sans-serif, system-ui, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
}
* { box-sizing: border-box; }
html, body { margin: 0; padding: 0; background: var(--bg); color: var(--text); font-family: var(--font); }
body { min-height: 100vh; }
a { color: var(--accent); }
img { display: block; max-width: 100%; }

.site-header {
  position: sticky;
  top: 0;
  z-index: 20;
  backdrop-filter: blur(12px);
  background: rgba(246, 244, 238, 0.9);
  border-bottom: 1px solid rgba(93, 77, 31, 0.12);
}
.site-header__inner {
  max-width: var(--max);
  margin: 0 auto;
  padding: 1rem 1.2rem;
  display: grid;
  grid-template-columns: 1fr minmax(240px, 340px);
  gap: 1rem;
  align-items: end;
}
.eyebrow {
  margin: 0 0 0.35rem;
  letter-spacing: 0.08em;
  text-transform: uppercase;
  color: var(--muted);
  font-size: 0.78rem;
}
.site-title {
  margin: 0;
  font-size: clamp(1.7rem, 3vw, 2.4rem);
}
.site-subtitle {
  margin: 0.4rem 0 0;
  color: var(--muted);
  max-width: 70ch;
}
.search-label {
  display: block;
  margin-bottom: 0.35rem;
  font-size: 0.9rem;
  color: var(--muted);
}
.search-input {
  width: 100%;
  border: 1px solid var(--border);
  background: white;
  border-radius: 999px;
  padding: 0.85rem 1rem;
  font: inherit;
  box-shadow: inset 0 1px 0 rgba(0,0,0,0.03);
}

.layout {
  max-width: var(--max);
  margin: 0 auto;
  padding: 1.2rem;
  display: grid;
  grid-template-columns: minmax(240px, 320px) minmax(0, 1fr);
  gap: 1.2rem;
  align-items: start;
}
.sidebar {
  position: sticky;
  top: 106px;
}
.sidebar-card,
.panel,
.info-card,
.sheet-card {
  background: var(--panel);
  border: 1px solid var(--border);
  box-shadow: var(--shadow);
  border-radius: var(--radius);
}
.sidebar-card {
  padding: 1rem 1rem 0.85rem;
  margin-bottom: 0.9rem;
}
.sidebar-title {
  margin: 0;
  font-weight: 700;
}
.sidebar-count {
  margin: 0.35rem 0 0;
  color: var(--muted);
}
.species-list {
  display: grid;
  gap: 0.7rem;
}
.species-list button {
  width: 100%;
  padding: 0;
  border: 0;
  background: none;
  text-align: left;
  cursor: pointer;
}
.species-pill {
  display: grid;
  grid-template-columns: 72px minmax(0, 1fr);
  gap: 0.85rem;
  align-items: center;
  padding: 0.7rem;
  border-radius: var(--radius);
  border: 1px solid var(--border);
  background: var(--panel);
  transition: transform 120ms ease, box-shadow 120ms ease, border-color 120ms ease;
  box-shadow: var(--shadow);
}
.species-pill:hover {
  transform: translateY(-1px);
  border-color: rgba(92, 75, 31, 0.4);
}
.species-pill.is-active {
  outline: 2px solid rgba(92, 75, 31, 0.18);
  border-color: rgba(92, 75, 31, 0.42);
  background: #fff8ea;
}
.species-pill__thumb {
  width: 72px;
  height: 72px;
  border-radius: 12px;
  overflow: hidden;
  background: var(--panel-strong);
}
.species-pill__thumb img {
  width: 100%;
  height: 100%;
  object-fit: cover;
}
.species-pill__name {
  font-weight: 700;
  line-height: 1.15;
}
.species-pill__meta {
  margin-top: 0.28rem;
  color: var(--muted);
  font-size: 0.92rem;
}

.content {
  min-width: 0;
}
.empty-state {
  padding: 2rem;
  border-radius: var(--radius);
  background: var(--panel);
  border: 1px solid var(--border);
  box-shadow: var(--shadow);
}
.sheet {
  display: grid;
  gap: 1rem;
}
.hero {
  display: grid;
  grid-template-columns: minmax(0, 1.1fr) minmax(320px, 0.9fr);
  gap: 1rem;
}
.hero-copy,
.hero-media,
.section-grid,
.two-up {
  min-width: 0;
}
.panel {
  padding: 1rem;
}
.hero-copy {
  display: grid;
  gap: 1rem;
}
.hero-title {
  margin: 0;
  font-size: clamp(2rem, 4vw, 3rem);
  line-height: 1.02;
}
.hero-scientific {
  margin: 0.3rem 0 0;
  font-size: 1.05rem;
  color: var(--muted);
}
.hero-scientific em { font-style: italic; }
.hero-lede {
  margin: 0;
  font-size: 1.03rem;
  line-height: 1.6;
}
.byline {
  color: var(--muted);
  font-size: 0.96rem;
}
.quick-facts {
  display: grid;
  grid-template-columns: repeat(auto-fit, minmax(160px, 1fr));
  gap: 0.7rem;
}
.fact-chip {
  background: var(--panel-strong);
  padding: 0.8rem;
  border-radius: var(--radius-small);
  border: 1px solid rgba(0,0,0,0.05);
}
.fact-chip__label {
  display: block;
  font-size: 0.8rem;
  color: var(--muted);
  margin-bottom: 0.25rem;
}
.fact-chip__value {
  font-weight: 700;
  line-height: 1.3;
}
.hero-media {
  display: grid;
  gap: 0.8rem;
}
.main-image-wrap {
  overflow: hidden;
  border-radius: var(--radius);
  background: var(--panel-strong);
  border: 1px solid var(--border);
  aspect-ratio: 4 / 3;
}
.main-image {
  width: 100%;
  height: 100%;
  object-fit: cover;
}
.thumbnail-row {
  display: grid;
  grid-template-columns: repeat(auto-fit, minmax(86px, 1fr));
  gap: 0.65rem;
}
.thumbnail-button {
  border: 1px solid var(--border);
  border-radius: 12px;
  overflow: hidden;
  background: white;
  padding: 0;
  cursor: pointer;
}
.thumbnail-button.is-active {
  outline: 2px solid rgba(92, 75, 31, 0.22);
  border-color: rgba(92, 75, 31, 0.48);
}
.thumbnail-button img {
  width: 100%;
  aspect-ratio: 1 / 1;
  object-fit: cover;
}
.hero-caption {
  color: var(--muted);
  font-size: 0.95rem;
  line-height: 1.5;
}
.section-title {
  margin: 0 0 0.6rem;
  font-size: 1.28rem;
}
.section-text,
.muted {
  color: var(--muted);
  line-height: 1.65;
}
.two-up {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: 1rem;
}
.list {
  margin: 0.25rem 0 0;
  padding-left: 1.1rem;
  line-height: 1.65;
}
.list li + li { margin-top: 0.4rem; }
.meta-grid {
  display: grid;
  gap: 0.5rem;
}
.meta-row {
  display: grid;
  grid-template-columns: 180px minmax(0, 1fr);
  gap: 0.75rem;
  padding: 0.45rem 0;
  border-bottom: 1px dashed rgba(0,0,0,0.08);
}
.meta-row:last-child { border-bottom: 0; }
.meta-label { color: var(--muted); }
.badge-row {
  display: flex;
  flex-wrap: wrap;
  gap: 0.55rem;
}
.badge {
  display: inline-flex;
  align-items: center;
  gap: 0.35rem;
  padding: 0.4rem 0.65rem;
  background: #fff8ea;
  border: 1px solid var(--accent-soft);
  border-radius: 999px;
  font-size: 0.9rem;
}
.map-grid {
  display: grid;
  grid-template-columns: minmax(0, 1.4fr) minmax(280px, 0.7fr);
  gap: 1rem;
  align-items: stretch;
}
.map-shell {
  overflow: hidden;
  min-height: 420px;
}
.map {
  width: 100%;
  min-height: 420px;
  border-radius: var(--radius-small);
}
.map-notes {
  display: grid;
  gap: 0.8rem;
}
.source-list,
.literature-list {
  display: grid;
  gap: 0.8rem;
}
.source-item,
.literature-item {
  padding: 0.9rem;
  background: var(--panel-strong);
  border: 1px solid rgba(0,0,0,0.06);
  border-radius: var(--radius-small);
}
.source-item__label,
.literature-item__title {
  font-weight: 700;
}
.source-item__meta,
.literature-item__meta {
  color: var(--muted);
  margin-top: 0.3rem;
  line-height: 1.55;
}
.note-box {
  padding: 0.9rem;
  border-radius: var(--radius-small);
  background: #fff8ea;
  border: 1px solid var(--accent-soft);
}
.footer-note {
  color: var(--muted);
  font-size: 0.92rem;
}
.hidden { display: none; }

@media (max-width: 1100px) {
  .layout { grid-template-columns: 1fr; }
  .sidebar { position: static; }
}
@media (max-width: 900px) {
  .site-header__inner,
  .hero,
  .two-up,
  .map-grid { grid-template-columns: 1fr; }
}
@media (max-width: 640px) {
  .layout,
  .site-header__inner { padding-left: 0.9rem; padding-right: 0.9rem; }
  .meta-row { grid-template-columns: 1fr; gap: 0.2rem; }
}